In addition to conventional galaxies, the universe contains very dim galaxies that until recently went unnoticed by astronomers. Possibly as numerous as conventional galaxies, these galaxies have the same general shape and even the same approximate number of stars as a common type of conventional galaxy, the spiral, but tend to be much larger. Because these galaxies' mass is spread out over larger areas, they have far fewer stars per unit volume than do conventional galaxies. Apparently these low-surface-brightness galaxies, as they are called, take much longer than conventional galaxies to condense their primordial gas and convert it to stars—that is, they evolve much more slowly.

    These galaxies may constitute an answer to the longstanding puzzle of the missing baryonic mass in the universe. Baryons—subatomic particles that are generally protons or neutrons—are the source of stellar, and therefore galactic, luminosity, and so their numbers can be estimated based on how luminous galaxies are. However, the amount of helium in the universe, as measured by spectroscopy, suggests that there are far more baryons in the universe than estimates based on galactic luminosity indicate. Astronomers have long speculated that the missing baryonic mass might eventually be discovered in intergalactic space or as some large population of galaxies that are difficult to detect.

    Anthropologists once thought that the ancestors of modern humans began to walk upright because it freed their hands to use stone tools ,which they had begum to make as the species evolved a brain of increased size and mental capacity. But discoveries of the three-million-year-old fossilized remains of our hominid ancestor Australopithecus have yielded substantial anatomical evidence that upright walking Appeared prior to the dramatic enlargement of the brain and the development of stone tools. Walking on two legs in an upright posture (bipedal locomotion) is a less efficient proposition than walking on all fours (quadrupedal locomotion) because several muscle groups that the quadruped uses for propulsion must instead to provide the biped the biped with stability and control. The shape and configuration of various bones must likewise be modified to allow the muscles to perform these functions in upright walking. Reconstruction of the pelvis (hipbones) and femur (thighbone) of "Lucy", a three-million-year-old skeleton that is the most complete fossilized skeleton from the australopithecine era, has shown that they are much more like he corresponding bones of the modern human than like those of the most closely related living primate, the quadrupedal chimpanzee. Lucy's wide, shallow pelvis is actually better suited to bipedal walking than is the rounder, bowl-like pelvis of the modern human, which evolved to form the larger birth canal needed to accommodate the head of a largebrained human infant .By contrast, the bead of Lucy's baby could have been no larger than that of a baby chimpanzee.

    If the small-brained australopithecines were not toolmakers, what evolutionary advantage did they gain by walking upright? One theory is that bipedality evolved in conjunction with the nuclear family: monogamous parents cooperating to care for their offspring. Walking upright permitted the father to use his hands to gather food and carry it to his mate from a distance, allowing the mother to devote more time and energy to nurturing and protecting their children. According to this view, the transition to bipedal walking may have occurred as long as ten million years ago, at the time of the earliest hominids, making it a crucial initiating event in human evolution.